Flora of China
  FOC Vol.2-3 (2013)              | Family List  Woodsiaceae  PDF     | Link Font:+ Big | - Small
   27. Woodsiaceae  
岩蕨科   yan jue ke
Authors:Authors: Zhang Gangmin, Masahiro Kato & Alexandr Shmakov


Plants epilithic, sometimes xerophytic (growing in dry habitats), small or medium-sized. Rhizomes short, erect, decumbent, or ascending, simple dictyostelic, covered with scales. Fronds clustered, monomorphic, deciduous or sometimes evergreen; stipe mostly shorter than lamina, ± scaly and hairy, articulate or not articulate; lamina 1-pinnate to bipinnatifid, elliptic-lanceolate to narrowly lanceolate, herbaceous or papery, frequently covered with articulate multicellular hairs, sometimes with glandular hairs or capitate glands. Veins free, pinnate, usually ending in enlarged hydathodes. Sori orbicular, consisting of 3-18 sporangia, borne at middle or apex of veinlets; indusia inferior, saucer-shaped to cup-shaped, margin long ciliate, or indusia absent, degenerated into curly, multicellular hairs (Woodsia), or sphaeropteroid (bladderlike) with openings at tip (Protowoodsia), sometimes also covered with false indusia, i.e., reflexed leaf margins (Cheilanthopsis). Spores ellipsoid or somewhat spherical, monolete, wall surface folded, cristate, tuberculate, or echinate.

About four genera (including the Caucasian Hymenocystis C. A. Meyer) and ca. 43 species: widely distributed in the N temperate and frigid zones, rarely in Central and South America, Africa (Angola, South Africa), and Madagascar; three genera and 24 species (nine endemic) in China.

The current classification of the genera requires evaluation by molecular phylogenetic evidence.

Wu Shiew-hung. 1999. Woodsiaceae. In: Wu Shiew-hung, ed., Fl. Reipubl. Popularis Sin. 4(2): 166-191.

1Stipe obviously articulate; indusia saucer- to cup-shaped, margin long ciliate.3  Woodsia    岩蕨属
+Stipe continuous; indusia spherical or degenerated into curly, multicellular hairs(2)
2(1)Indusia platelike with unequal lobes or dissected with filiform lobes.3  Woodsia    岩蕨属
+Indusia spherical, tearing when mature(3)
3(2)Lamina glabrous or with few glandular trichomes, margin flat, not reflexed; pinnae not articulate to rachis.1  Protowoodsia    膀胱蕨属
+Lamina pubescent, margin reflexed to form false indusia (usually fallen later); pinnae articulate to rachis.2  Cheilanthopsis    滇蕨属
   Lower Taxon
  • Cheilanthopsis  Hieronymus  滇蕨属
  • Protowoodsia  Ching  膀胱蕨属
  • Woodsia  R. Brown  岩蕨属