Flora of China
  FOC Vol.15 (1996)              | Family List  Styracaceae  PDF     | Link Font:+ Big | - Small
   Styracaceae  Dumortier
安息香科   an xi xiang ke
Authors:Hwang Shu-mei; James Grimes


Trees or shrubs, usually stellate pubescent or scaly, rarely glabrous. Leaves usually alternate, simple; stipules absent or very minute. Inflorescences terminal or axillary, racemes, panicles, or cymes, rarely 1-flowered or several flowers in a fascicle; bracteoles minute or absent. Flowers bisexual, rarely polygamous dioecious, actinomorphic. Calyx campanulate, obconical, or cup-shaped; tube completely or partially adnate to ovary; teeth or lobes 4 or 5(or 6), sometimes very small or obsolete. Corolla mostly white, gamopetalous; lobes (4 or)5(--7), basally ± connate, rarely free, imbricate or valvate, rarely slightly induplicate. Stamens twice, sometimes equal in number to corolla lobes, inserted at base of corolla; filaments mostly flattened, basally partially or completely connate into a tube; anthers introrse, 2-locular, locules parallel and opening by longitudinal slits. Ovary superior, half inferior, or inferior, 3--5-locular or apically 1-locular and basally 3--5-locular; ovules few or solitary in each locule, erect, pendulous, or anatropous, integument 1 or 2, placentation axile or parietal. Style slender, linear or subulate; stigma truncate, capitate or 2--5-lobed. Fruit a berry, drupe, or capsule, exocarp fleshy to dry. Seeds sometimes winged, often with a broad hilum; embryo straight or slightly curved; endosperm copious; cotyledons flattened or subterete.

Eleven genera and ca. 180 species: tropical and temperate America, Asia, and Mediterranean; 10 genera (two endemic) and 54 species (32 endemic) in China.

Hwang Shu-mei in Wu Rong-fen (as Wu Young-fen) & Hwang Shu-mei, eds. 1987. Styracaceae. Fl. Reipubl. Popularis Sin. 60(2): 77-150.

1Fruit completely separate from or slightly adnate to persistent calyx; ovary superior.(2)
+Fruit partially or almost completely adnate to persistent calyx; ovary inferior.(4)
2(1)Ovary 3-locular when young, becoming 1-locular; seeds 1 or 2, without wings, rounded at both ends; pedicel not jointed1  Styrax    安息香属
+Ovary 5(or 6)-locular; seeds numerous, winged or acute at both ends; pedicel jointed.(3)
3(2)Filaments coherent or slightly connate at base; fruit drupaceous, indehiscent, pear-shaped; seeds pointed at both ends2  Bruinsmia    歧序安息香属
+Filaments connate at least to middle; fruit capsular dehiscing loculicidally by 5 valves; seeds winged at both ends3  Alniphyllum    赤杨叶属
4(1)Fruit dehiscing loculicidally by 3 valves; seeds winged at both ends; petals free, connective 2- or 3-dentate4  Huodendron    山茉莉属
+Fruit indehiscent; seeds without wings; petals connate at base (shortly coherent in Pterostyrax), connective entire.(5)
5(4)Corolla lobes 4(or 5); fruit with 2--4 wings5  Halesia    银钟花属
+Corolla lobes 5 (4--7 in Sinojackia); fruit with 5--12 ribs or wings (Parastyrax without ribs or wings).(6)
6(5)Flowers solitary or 2 together; filaments of equal length6  Melliodendron    陀螺果属
+Flowers in racemes, panicles, or thyrses; filaments 5 long and 5 short or subequal (sometimes equal in Sinojackia).(7)
7(6)Leaf blade margin entire or callose-denticulate; inflorescences crowded10  Parastyrax    茉莉果属
+Leaf blade margin serrate or serrulate; inflorescences not crowded.(8)
8(7)Winter buds enclosed by scales; fruit apex with a short point9  Rehderodendron    木瓜红属
+Winter buds naked; fruit apex tapering into a beak.(9)
9(8)Inflorescences paniculate; calyx tube campanulate; fruit exocarp thin, crusty7  Pterostyrax    白辛树属
+Inflorescences racemose cymes; calyx tube obconical; fruit exocarp thick, woody or fleshy8  Sinojackia    秤锤树属
   Lower Taxon
  • Alniphyllum  Matsumura  赤杨叶属
  • Bruinsmia  Boerlage & Koorders  歧序安息香属
  • Halesia  J. Ellis ex Linnaeus  银钟花属
  • Huodendron  Rehder  山茉莉属
  • Melliodendron  Handel-Mazzetti  陀螺果属
  • Parastyrax  W. W. Smith  茉莉果属
  • Pterostyrax  Siebold & Zuccarini  白辛树属
  • Rehderodendron  Hu  木瓜红属
  • Sinojackia  Hu  秤锤树属
  • Styrax  Linnaeus  安息香属