Flora of China
  FOC Vol.4 (1999)              | Family List  Podocarpaceae  PDF     | Link Font:+ Big | - Small
   Podocarpaceae  Endlicher
罗汉松科   luo han song ke
Authors:Fu Liguo (傅立国 Fu Li-kuo) , Li Yong (李勇) ; Robert R. Mill


Trees or shrubs evergreen, dioecious or rarely monoecious. Leaves decussate, subopposite, or spirally arranged; blade scalelike, subulate, or linear to elliptic, stomatal lines abaxial or present on all surfaces. Pollen cones terminal, solitary or clustered in leaf axils, or borne in spikelike complexes; individual cones pedunculate or sessile; microsporophylls numerous, spirally arranged, with distinct adaxial and abaxial surfaces; microsporangia 2; pollen 2(or 3)-saccate in Chinese species, (rarely nonsaccate). Seed-bearing structures terminal or axillary, solitary, occasionally spikelike, comprising few to several spirally arranged bracts; all or only apical bracts fertile, smooth or warty; basal bracts sometimes fused and succulent (together with peduncle) to form a "receptacle," or obsolete; ovule (inverted) or inclined in Chinese species. Seed drupelike or nutlike, wholly or (in Dacrydium) partly enveloped in a sometimes colored and succulent epimatium derived from fertile ovulate scale. Cotyledons 2.

Eighteen genera and ca. 180 species: tropical, subtropical, and S temperate zones, mainly in S hemisphere but extending to montane tropical Africa, Central America, and Japan; four genera and 12 species (three endemic) in China.

Although strictly a flowering plant term, "receptacle" is used widely in Podocarpaceae literature, e.g., by Tomlinson (Int. J. Plant Sci. 153: 572-588. 1992). Spjut (Mem. New York Bot. Gard. 70: 21, 71-73. 1994) incorrectly used the term "epimatium" to denote the peduncle of Nageia and also the receptacle of Podocarpus and Dacrycarpus.

Cheng Wan-chün, Fu Li-kuo & Chao Chi-son. 1978. Podocarpaceae. In: Cheng Wan-chün & Fu Li-kuo, eds., Fl. Reipubl. Popularis Sin. 7: 398-422.

1Leaves dimorphic: juvenile leaves needlelike or linear; adult leaves needlelike or scalelike, less than 5 mm.(2)
+Leaves ± monomorphic: adult leaves neither needlelike nor scalelike, more than 5 mm; juvenile leaves similar to adult leaves in shape but often larger and/or wider.(3)
2(1)Juvenile leaves 2-ranked, arranged in 1 plane (forming an oblong-ovate branchlet outline), linear, not scalelike; adult leaves needlelike or scalelike, falcate, 0.8-1.5 mm2  Dacrycarpus    鸡毛松属
+Juvenile leaves not 2-ranked, spreading, not arranged in 1 plane, linear to needlelike or subulate; adult leaves hard and scalelike, 2-5 mm1  Dacrydium    陆均松属
3(1)Leaf blade with a single, obvious, often raised midvein on 1 or both surfaces4  Podocarpus    罗汉松属
+Leaf blade without an obvious midvein but with many, slender, longitudinal veins dichotomous in basal region and converging toward apex3  Nageia    竹柏属
   Lower Taxon
  • Dacrycarpus  (Endlicher) de Laubenfels  鸡毛松属
  • Dacrydium  Solander ex G. Forster  陆均松属
  • Nageia  Gaertner  竹柏属
  • Podocarpus  L’Heritier ex Persoon  罗汉松属