Flora of China
  FOC Vol.2-3 (2013)              | Family List  Marattiaceae  PDF     | Link Font:+ Big | - Small
   8. Marattiaceae  
合囊蕨科   he nang jue ke
Authors:Authors: He Zhaorong & Maarten J. M. Christenhusz


Plants terrestrial (rarely epiphytic, sometimes in streams), evergreen. Rhizomes erect, ascending, or creeping, amylaceous (starchy), polycyclostelic to dictyostelic. Fronds monomorphic and long-lived or dimorphic and fertile ones short-lived (Danaea), simple or compound, 1-4-pinnate or (pedate-)palmate (Christensenia), small (ca. 20 cm) to very large (up to at least 6 m). Stipes fleshy, with a pair of stipulelike appendages at base, often with conspicuous lenticels. Pulvini at base of fronds, at nodes, at bases of pinnae, and in some species at “naked” nodes (without pinnae) along stipe. Veins free, simple or bifurcate, or reticulate (Christensenia), false veins present between true veins or absent. Tissues mucilaginous. Indument of multicellular uniseriate hairs and basifixed or peltate scales. Sori lacking true indusia, usually with paraphyses, superficial, sunken or stalked, elongate along veins, bilateral or radial. Sporangia partially or completely fused into synangia, dehiscing by a slit or pore. Spores trilete or monolete, with bullate, muriform, or stellate ornamentation. Prothalli very large, thalloid, photosynthetic.

Six genera and ca. 100 species: pantropical, Danaea Smith, Eupodium J. Smith, and Marattia Swartz in the Neotropics, Angiopteris, Christensenia, and Ptisana in the Paleotropics; three genera and 30 species (17 endemic) in China.

The delimitation of certain species of Angiopteris is unclear.

The following papers are relevant: Christenhusz (Ann. Univ. Turku., A II, 216: 1-134. 2007); Christenhusz and Toivonen (Biol. Invas. 10: 1215-1228. 2008); Murdock (Amer. J. Bot. 95: 626-641. 2008); and Murdock (Taxon 57: 737-755. 2008).

Ching Ren-chang, Fu Shu-hsia, Wang Chu-hao & Shing Gung-hsia. 1959. Angiopteridaceae and Christenseniaceae. In: Ching Ren-chang, ed., Fl. Reipubl. Popularis Sin. 2: 27-66, 330-345; Zhang Xianchun. 2004. Marattiaceae. In: Zhang Xianchun, ed., Fl. Reipubl. Popularis Sin. 6(3): 246-248.

1Fronds simple, pedately lobed to palmately compound; venation reticulate; synangium radial.2  Christensenia    天星蕨属
+Fronds pinnate (sometimes simple when plants juvenile); venation free, simple or bifurcate; synangium ovate(2)
2(1)Sporangia fully fused into synangia, synangia bivalved.1  Ptisana    合囊蕨属
+Sporangia fused basally but free apically, synangia composed of globose sporangia.3  Angiopteris    莲座蕨属
   Lower Taxon
  • Angiopteris  Hoffmann  莲座蕨属
  • Christensenia  Maxon  天星蕨属
  • Ptisana  Murdock  合囊蕨属