Flora of China
  FOC Vol.6 (2001)              | Family List  Caryophyllaceae  PDF Arenaria  PDF   | Link Font:+ Big | - Small
   19. Arenaria  Linnaeus
无心菜属   wu xin cai shu
Authors:Wu Zhengyi; Warren L. Wagner


Herbs annual or perennial, rarely biennial. Stems erect or rarely creeping, often caespitose or pulvinate. Leaves opposite, rarely whorled; leaf blade linear to elliptic, ovate, or orbicular, usually flat, margin entire. Flowers solitary or numerous in cymes, actinomorphic. Sepals 4 or 5, apex entire, rarely emarginate. Petals 4 or 5, sometimes absent, apex entire to toothed, 2-cleft, or fimbriate. Stamens (2--5 or 8 or)10. Ovary 1-loculed; ovules numerous. Styles 2 or 3(--5). Capsule ovoid, obovoid, or globose, usually shorter than persistent sepals, rarely equaling or longer than them, 3- or 6-valved. Seeds reniform or subovoid, flattened, smooth, tuberculate, or narrowly winged.

More than 300 species: N temperate to arctic regions; 102 species (79 endemic) in China.

Arenaria has been subdivided into ten subgenera, several of which were further divided into sections and series (McNeill, Notes Roy. Bot. Gard. Edinburgh 24: 79–155. 1962). Six subgenera are represented in China, one of which ( A. subgen. Odontostemma ) contains more than half the species.

Arenaria ladyginii Kozhevnikov (Novosti Sist. Vyssh. Rast. 21: 67. 1984) was described from Xizang. However, the type has not been seen by the present authors and its identity could not be ascertained.

1Styles usually 3 and capsules 6-valved or -toothed, or styles rarely 2 and capsules 4-valved or -toothed; petal apex entire or slightly emarginate; sepals never truncate and saccate at base, not excurved.(2)
+Styles usually 2, sometimes 3(or 4 or 5); capsule 4-valved or -toothed, rarely 6-, 8-, or 10-valved or -toothed; petal apex entire to fimbriate; sepals truncate and ± saccate at base, excurved (6. A. subgen. Odontostemma).(44)
2(1)Sepals not incrassate at base, veins not obviously impressed; leaf blade subulate to ovate, margin and apex incrassate, apex not setiform but spinose.(3)
+Sepals usually (rarely not) incrassate at base, veins usually obvious (rarely not); leaf blade variously shaped, margin and apex not incrassate, or margin scarious, apex spinose when blade aristate.(13)
3(2)Sepal apex obtuse, margin scarious; basal leaf blade subulate to linear-lanceolate, cauline leaf blade imbricate in 4 rows (decussate), ovate, less than 5 mm, persistent; flowers solitary or 3 in cymes; plants congested caespitose or pulvinate (4. A. subg(4)
+Sepal apex acute, acuminate, or obtuse, apex and margin usually hard; leaf blade remote or sometimes approximate (covering each other), but never imbricate in 4 rows, lanceolate, elliptic, ovate, or orbicular, more than 5 mm, deciduous, rarely persistent;(8)
4(3)Plants densely caespitose; stems 10--20 cm, glandular villous33  A. przewalskii    福禄草
+Plants densely pulvinate; stems 2.5--15 cm, glandular villous or glabrous.(5)
5(4)Stems 4--15 cm, villous or glandular hairy.(6)
+Stems 2.5--6 cm, glabrous.(7)
6(5)Stems 10--15 cm, villous; sepals ovate to lanceolate-ovate34  A. smithiana    大花福禄草
+Stems 4--9 cm, glandular villous; sepals elliptic35  A. oreophila    山生福禄草
7(5)Pedicels 2--4 cm; petals ca. 2 × as long as sepals; anthers violet36  A. densissima    密生福禄草
+Pedicels absent (flowers sessile); petals slightly longer than sepals; anthers yellow37  A. polytrichoides    团状福禄草
8(3)Floral disc large, with 5 obvious glands.(9)
+Floral disc small or not impressed, glands small or degenerate.(10)
9(8)Stems 1--15 cm, villous on one side; leaf blade glabrous except ciliate at base; petals white or pink42  A. forrestii    西南无心菜
+Stems 2--6 cm, villous all round; leaf blade glandular pubescent; petals violet43  A. glanduligera    小腺无心菜
10(8)Petals violet; anthers violet41  A. rhodantha    红花无心菜
+Petals white or pink; anthers pale yellow, green, or black.(11)
11(10)Plants hairy throughout; stems diffuse; leaf blade ovate or ovate-orbicular, margin long hairy; petals white; anthers dark green or black38  A. ciliolata    缘毛无心菜
+Plants not hairy throughout; stems erect; leaf blade lanceolate or narrowly orbicular, margin shortly hairy or glabrous; petals white or pink; anthers yellow or green.(12)
12(11)Stems 4--6 cm; leaf blade lanceolate or ovate-lanceolate, margin shortly hairy; inflorescence rachis bracteate proximally; sepal apex excurved; petals white or pink; anthers pale yellow39  A. ramellata    嫩枝无心菜
+Stems 2--5 cm; leaf blade narrowly orbicular, margin glabrous; inflorescence rachis ebracteate proximally; sepal apex erect; petals white; anthers green40  A. stracheyi    藏西无心菜
13(2)Leaf blade linear to orbicular, or aristate and narrowly linear, margin not obviously scarious; sepal veins inconspicuous; plants solitary or caespitose, rarely compacted pulvinate; leaves deciduous, rarely persistent.(14)
+Leaf blade subulate to linear, never setiform or graminoid, margin obviously scarious; sepal veins impressed; plants caespitose to compacted pulvinate; leaves persistent (3. A. subgen. Eremogoneastrum).(28)
14(13)Sepals slightly or not incrassate at base; leaf blade usually linear to ovate, rarely needle-shaped (1. A. subgen. Arenaria).(15)
+Sepals markedly incrassate or woody at base; leaf blade linear or linear-subulate, graminoid, rarely aristate with a spinose tip (2. A. subgen. Eremogone).(20)
15(14)Seeds with a flat wing; petals slightly shorter than or equaling sepals6  A. compressa    扁翅无心菜
+Seeds wingless; petals shorter than bracts, slightly shorter to longer than sepals.(16)
16(15)Seeds tuberculate with raised papillae; petals usually slightly shorter than sepals1  A. serpyllifolia    无心菜
+Seeds not tuberculate with raised papillae; petals equaling or longer than sepals.(17)
17(16)Seeds reticulate; inflorescence of terminal cymes; leaf blade linear to ovate-saccate.(19)
+Seeds with curls; inflorescence usually axillary; leaf blade elliptic to ovate or orbicular.(18)
18(17)Leaf blade elliptic, ovate, or suborbicular; sepals glabrous abaxially2  A. orbiculata    圆叶无心菜
+Leaf blade elliptic-lanceolate or obovate; sepals sparsely white villous on veins abaxially3  A. neelgherrensis    尼盖无心菜
19(17)Capsule 1-seeded; leaf blade linear; sepals with papillose spots abaxially4  A. monosperma    单子无心菜
+Capsule several seeded; leaf blade elliptic to elliptic-ovate; sepals white glandular villous abaxially5  A. puranensis    普兰无心菜
20(14)Plants completely woody at base; sepals not herbaceous, veins inconspicuous, apex usually acuminate.(21)
+Plants not completely woody at base; sepals herbaceous throughout or only at middle, veins impressed, apex usually obtuse, rarely acute or acuminate.(22)
21(20)Leaf blade apex acute; pedicels glabrous, rarely sparsely glandular hairy14  A. griffithii    裸茎老牛筋
+Leaf blade apex spinose; pedicels densely glandular hairy15  A. androsacea    点地梅状老牛筋
22(20)Sepals herbaceous only at middle, apex long acuminate, with (1--)3 or more parallel veins7  A. potaninii    五蕊老牛筋
+Sepals herbaceous ± throughout, apex obtuse, 1-veined.(23)
23(22)Stems with congested, pale brown, long, hardened, dead leaves at base; leaves forming a sheath at base8  A. juncea    老牛筋
+Stems without basal leaves as above; leaves not forming a sheath at base.(24)
24(23)Plants pulvinate; stems 4--10 cm; leaf blade linear to subulate, 1.5--5 cm.(25)
+Plants congested caespitose; stems 4--20 cm; leaf blade linear, 2--6 cm.(26)
25(24)Stems 4--10 cm, white glandular hairy distally; sepals ± glandular villous12  A. formosa    美丽老牛筋
+Stems 6--10 cm, villous distally; sepals glabrous13  A. haitzeshanensis    海子山老牛筋
26(24)Stems 4--10 cm, glandular pubescent distally11  A. grueningiana    华北老牛筋
+Stems 10--20 cm; all or only peduncle proximally glandular hairy, or glabrous.(27)
27(26)Stems glabrous proximally, becoming glandular hairy or villous distally; sepals glabrous9  A. capillaris    毛叶老牛筋
+Stems hirsutulous throughout; sepals densely glandular hairy10  A. acicularis    针叶老牛筋
28(13)Flowers few, in cymes.(29)
+Flowers solitary, rarely paired.(30)
29(28)Stems sparsely villous, 2--3 cm; leaf blade 5--6 mm; sepals 3-veined31  A. taibaishanensis    太白雪灵芝
+Stems glandular villous, to 8 cm; leaf blade 0.7--20 mm; sepals usually 1-veined32  A. festucoides    狐茅状雪灵芝
30(28)Flowers sessile.(31)
+Flowers pedicellate.(34)
31(30)Sepals violet abaxially; petals pink, lanceolate or orbicular27  A. shannanensis    粉花雪灵芝
+Sepals green; petals white, ovate, obovate, or saccate.(32)
32(31)Flowers ca. 1.8 cm in diam.; sepals ovate-lanceolate; petals broadly obovate, ca. 1.5 × as long as sepals28  A. edgeworthiana    山居雪灵芝
+Flowers ca. 0.8 cm in diam.; sepals elliptic-lanceolate or lanceolate; petals narrowly obovate or saccate, shorter or slightly longer than sepals.(33)
33(32)Sepals elliptic-lanceolate; petals narrowly obovate, slightly longer than sepals; anthers equal; styles equaling ovary29  A. bryophylla    藓状雪灵芝
+Sepals lanceolate; petals saccate, shorter than sepals; anthers 5 long and 5 short; styles ca. 2.5 × as long as ovary30  A. zadoiensis    杂多雪灵芝
34(30)Pedicels glandular villous or villous.(35)
+Pedicels glabrous.(39)
35(34)Pedicels villous.(36)
+Pedicels glandular villous.(37)
36(35)Sepals lanceolate, glabrous19  A. kansuensis    甘肃雪灵芝
+Sepals narrowly lanceolate, sparsely pilose, at least at base20  A. ischnophylla    瘦叶雪灵芝
37(35)Sepals ovate-lanceolate, 3-veined; petals ovate, ca. 1/3--1/2 as long as sepals16  A. brevipetala    雪灵芝
+Sepals lanceolate or ovate-lanceolate, 1-veined; petals elliptic, slightly longer than sepals.(38)
38(37)Flowers solitary, rarely paired, pedicel 2.5--5 × as long as bracts; anthers yellow17  A. baxoiensis    八宿雪灵芝
+Flowers solitary, never paired, pedicel shorter than bracts; anthers black18  A. kumaonensis    库莽雪灵芝
39(34)Pedicels 4--10 mm.(40)
+Pedicels 1--2 mm.(41)
40(39)Pedicels 5--10 mm; sepals lanceolate, ca. 5 mm; anthers yellow; petals not callose at base21  A. roborowskii    青藏雪灵芝
+Pedicels ca. 4 mm; sepals elliptic, 3 mm; anthers white; petals with a yellow, oblong callus at base22  A. lancangensis    澜沧雪灵芝
41(39)Sepals hard, 1-veined, base ciliate, apex greenish brown23  A. gerzensis    改则雪灵芝
+Sepals soft, 1--3-veined, base not ciliate, apex green.(42)
42(41)Flowers 6--7 mm in diam.26  A. pulvinata    垫状雪灵芝
+Flowers ca. 15 mm in diam.(43)
43(42)Sepals linear-lanceolate, 7--8 mm, 1--3 veined; petals elliptic-ovate, 8--9 mm; anthers yellow24  A. qinghaiensis    青海雪灵芝
+Sepals lanceolate, 6--7 mm, closely 3-veined; petals obovate, 4--5 mm; anthers violet25  A. aksayqingensis    阿克赛钦雪灵芝
44(1)Petal apex toothed or fimbriate; filaments opposite sepals with a gland at base; styles 2.(45)
+Petal apex entire, slightly emarginate, or shallowly 2-cleft, rarely lobes with 3 shallow, round teeth; filaments opposite sepals with a gland at base or not; styles 2 or 3(--5).(67)
45(44)Petals fimbriate or toothed.(46)
+Petal apex toothed.(56)
46(45)Roots conic.(47)
+Roots fusiform, solitary or caespitose, rarely each node of rhizome globose or clavate, forming a moniliform chain.(48)
47(46)Leaf blade 0.5--1.5 × 0.2--1 cm; sepals 4--10 mm; petals slightly longer than sepals101  A. delavayi    大理无心菜
+Leaf blade 1--3 × 0.5--0.6 cm; sepals 3--6 mm; petals ca. 2 × as long as sepals102  A. yunnanensis    云南无心菜
48(46)Roots moniliform100  A. monilifera    念珠无心菜
+Roots fusiform.(49)
49(48)Leaves glabrous; sepal margin black ciliate92  A. salweenensis    怒江无心菜
+Leaves often glandular pubescent; sepals glandular pubescent but margin not black ciliate.(50)
50(49)Anthers yellow, rarely green.(51)
+Anthers black, violet, or yellowish brown (rarely yellow in A. iochanensis).(53)
51(50)Stems 10--25 cm; leaf blade narrowly elliptic, orbicular-elliptic, or orbicular-lanceolate; pedicels villous or glandular hairy97  A. fimbriata    瓣无心菜
+Stems 40--60 cm; leaf blade narrowly elliptic-saccate, ovate-elliptic, or ovate; pedicels multicellular pubescent or glandular hairy.(52)
52(51)Plants annual or biennial; stems 30--60 cm; leaves to 40 cm; seeds 2 or 398  A. debilis    柔软无心菜
+Plants perennial; stems 10--40 cm; leaves to 15 cm; seeds 11 or 1299  A. polysperma    多子无心菜
53(50)Plants hispid; leaf blade ovate or orbicular-elliptic93  A. trichophora    具毛无心菜
+Plants glandular or shortly villous; leaf blade broadly lanceolate, orbicular-ovate, or orbicular.(54)
54(53)Stem solitary, branched below middle; leaf blade orbicular-obovate or orbicular, 3--10 mm wide; seeds 2--4, wingless; roots several clustered together96  A. barbata    髯毛无心菜
+Stems branched at base, procumbent, diffuse, or erect; leaf blade lanceolate, 1--4 mm wide; seeds numerous, with a narrow wing; root solitary.(55)
55(54)Stems procumbent or erect; leaf blade usually broadly lanceolate, 2--4 mm wide; pedicels 0.5--1.5 cm; petal apex fimbriate; anthers violet or black94  A. yulongshanensis    狭叶无心菜
+Stems diffuse; leaf blade narrowly lanceolate, 1--3 mm wide; pedicels 0.5--2 cm; petal apex slightly toothed; anthers yellow95  A. iochanensis    药山无心菜
56(45)Roots solitary or several clustered together, spinose, rarely conic.(57)
+Roots conic, fleshy, rarely fibrous.(62)
57(56)Stems slender; leaves glabrous or sparsely hairy; petal apex 4-toothed.(58)
+Stems rigid; leaves variously hairy; petal apex with small teeth.(59)
58(57)Stems glandular pubescent with 2 lines of hairs; leaves sparsely hairy; petals obovate or narrowly elliptic80  A. quadridentata    四齿无心菜
+Stems and leaves glabrous; petals obovate or cuneate81  A. giraldii    秦岭无心菜
59(57)Leaf blade ovate or ovate-orbicular; sepals orbicular-ovate or orbicular-lanceolate85  A. fridericae    玉龙山无心菜
+Leaf blade mostly ovate-lanceolate or ovate; sepals lanceolate, rarely ovate.(60)
60(59)Sepals long villous; petals longer than sepals, but less than 2 × as long as them82  A. trichophylla    毛叶无心菜
+Sepals villous or glandular hairy; petals more than 1.5 × as long as sepals.(61)
61(60)Stems to 1.5 cm, with numerous branches; flower solitary, terminal, to 1.5 cm in diam.; anthers violet to yellow-brown83  A. pogonantha    须花无心菜
+Stems to 30 cm, distally with branched, dichasial cymes; flowers small, to 0.8 cm in diam.; anthers blue-green to yellow84  A. dimorphitricha    滇蜀无心菜
62(56)Roots fibrous; stems brown glandular villous; leaf blade tuberculate on both surfaces, petiole 2--5 mm86  A. longicaulis    长茎无心菜
+Roots conic, fleshy; stems glabrous, subglabrous, or glandular hairy; leaf blade not tuberculate, petiole 1--2 cm or leaves sessile.(63)
63(62)Stems glabrous or subglabrous; petiole 1--2 cm (at least proximal cauline leaves petiolate)87  A. longipetiolata    长柄无心菜
+Stems villous or glandular hairy; leaves sessile or shortly petiolate.(64)
64(63)Stems pale brown, villous.(65)
+Stems green, glandular hairy.(66)
65(64)Stems 4--7 cm; leaf blade elliptic or obovate-elliptic; sepals long villous abaxially88  A. leucasteria    毛萼无心菜
+Stems 10--15 cm; leaf blade elliptic to broadly elliptic-saccate; sepals sparsely glandular pubescent abaxially89  A. nivalomontana    大雪山无心菜
66(64)Leaf blade thin, glabrous or glandular hairy; sepals narrowly ovate, glabrous; petals ca. 1.5 × as long as sepals; cymes spreading90  A. omeiensis    峨眉无心菜
+Leaf blade thick, villous with yellowish, small, nodose hairs; sepals orbicular-lanceolate, long pubescent abaxially; petals slightly longer than sepals; cymes compact91  A. auricoma    黄毛无心菜
67(44)Filaments opposite sepals with a gland at base; seeds globose or reniform, smooth or tuberculate.(68)
+Filaments opposite sepals without a gland at base or gland not obvious; seeds oblate or reniform, glabrous or with small, raised tubercles, or flat but margin with a narrow wing.(72)
68(67)Stems villous with 2 lines of hairs; sepals glabrous or only ciliate at margin.(69)
+Stems glandular villous, or with long, nodose bristles; sepals glandular villous or glandular villous mixed with long, nodose bristles.(70)
69(68)Styles 6--7 mm; petal apex entire; leaves aggregated at stem apex, leaf blade broadly linear or linear-lanceolate75  A. longistyla    长柱无心菜
+Styles ca. 1.5 mm; petal apex 2-cleft; leaves in several distant pairs, leaf blade saccate76  A. ionandra    紫蕊无心菜
70(68)Stems and sepals densely hairy mixed with long, nodose bristles; petals ca. 10 mm77  A. longiseta    长刚毛无心菜
+Stems and sepals hairy, without nodose bristles; petals 5--8 mm.(71)
71(70)Leaf blade obsaccate, petiole short; sepals 2--3 mm78  A. chamdoensis    昌都无心菜
+Leaf blade linear-lanceolate, petiole absent; sepals 5--6 mm79  A. szechuensis    四川无心菜
72(67)Seeds usually flat, with a narrow wing.(73)
+Seeds reniform, subglobose, globose, or oblate, smooth or tuberculate, wingless.(82)
73(72)Stems diffuse; flowers usually several in cymes.(74)
+Stems erect or suberect; flowers solitary, rarely paired.(76)
74(73)Stems sparsely villous; petals oblanceolate or obovate, apex 2- or 3-toothed67  A. euodonta    真齿无心菜
+Stems glandular hairy; petals obovate or broadly ovate, apex entire or emarginate.(75)
75(74)Leaf blade orbicular-lanceolate or orbicular; petals broadly ovate, apex emarginate65  A. melanandra    黑蕊无心菜
+Leaf blade linear-lanceolate; petals ovate, apex entire66  A. pseudostellaria    线叶无心菜
76(73)Stems 3--6 cm, glabrous proximally68  A. dsharaensis    察龙无心菜
+Stems 4--25 cm, villous or glandular hairy.(77)
77(76)Plants to 6 cm tall, white villous in 2 lines or white, yellow, or black nodose villous; petals white.(78)
+Plants 4--25 cm tall, glandular villous; petals white, pink, or violet.(79)
78(77)Plants 5--6 cm tall; stems white villous in 2 lines; petal apex entire74  A. monantha    山地无心菜
+Plants to 4 cm tall; stems white, yellow, or black nodose villous; petal apex emarginate, often with a few small teeth73  A. inornata    无饰无心菜
79(77)Petal apex entire or emarginate; cauline leaf blade lanceolate, elliptic, ovate, or orbicular.(80)
+Petal apex entire or shallowly 2-cleft; cauline leaf blade linear-lanceolate or lanceolate, sometimes elliptic.(81)
80(79)Stems white; leaf blade violet; petal apex entire69  A. melandryoides    桃色无心菜
+Stems violet; leaf blade green; petal apex slightly emarginate70  A. rockii    紫红无心菜
81(79)Petals pink, apex shallowly 2-cleft71  A. roseiflora    粉花无心菜
+Petals white, apex entire72  A. melandryiformis    女娄无心菜
82(72)Plants usually 8--20 cm tall; proximal flowers of stem with petals and anthers reduced, distal flowers of stem with petals and anthers not reduced.(83)
+Plants rarely more than 8 cm tall, or very slender when more than 8 cm tall; petals and stamens sometimes degenerate or reduced to various degrees, petals (when present) ca. 2/3 as long as or equaling sepals.(90)
83(82)Styles 3(or 5).(84)
+Styles 2.(87)
84(83)Principal root napiform or conic, unbranched.(85)
+Principal root conic, fleshy, many branched.(86)
85(84)Petals white; stems white; leaves glabrous47  A. filipes    细柄无心菜
+Petals white or pink; stems white or violet; leaves ciliate at base, otherwise glabrous46  A. napuligera    滇藏无心菜
86(84)Petal apex shallowly 2-lobed, lobes with small, irregular teeth; stems and leaves with septate hairs45  A. setifera    刚毛无心菜
+Petal apex rounded, emarginate, or bifid to near middle of petal; stems and leaves with non-septate hairs44  A. weissiana    多柱无心菜
87(83)Leaf blade linear-lanceolate, to 1.5 cm, glabrous, margin not cartilaginous; stamens 551  A. schneideriana    雪山无心菜
+Leaf blade lanceolate to orbicular, to 4 cm, subglabrous, margin cartilaginous; stamens 10.(88)
88(87)Leaf blade narrowly elliptic to suborbicular, 8--15 mm wide48  A. nigricans    变黑无心菜
+Leaf blade lanceolate to oblanceolate, 1--5 mm wide.(89)
89(88)Leaf blade lanceolate or oblanceolate, 1--2 cm × 2--5 mm; sepals lanceolate, pubescent or with white, septate, glandular hairs, margin hairy; petal apex 2-cleft49  A. zhongdianensis    中甸无心菜
+Leaf blade oblanceolate, 0.5--2.5 cm × 1--5 mm; sepals ovate or ovate-lanceolate, densely shortly hairy at base; petal apex entire50  A. xerophila    旱生无心菜
90(82)Petals absent or only 2 or 3 developed, reduced to various degrees.(91)
+All floral parts present.(94)
91(90)Sepals 5; petals absent.(92)
+Sepals 4; petals present or absent.(93)
92(91)Stamens 5; leaf blade rhomboid-elliptic to saccate; stem with 1 or 2 lines of hairs61  A. reducta    减缩无心菜
+Stamens 2 or 3; leaf blade linear to linear-lanceolate; stem with 2 lines of hairs62  A. pharensis    帕里无心菜
93(91)Petals absent; stamens 2 or 3; sepals and leaves fleshy; pedicels and sepals glabrous63  A. littledalei    古临无心菜
+Petals present; stamens 8; sepals and leaves not fleshy; pedicels and sepals hairy64  A. saginoides    漆姑无心菜
94(90)Styles 3.(95)
+Styles 2(or 3).(96)
95(94)Leaf blade subulate, 4--5 × ca. 1 mm52  A. minima    微无心菜
+Leaf blade saccate, 5--12 × 2--3 mm53  A. bomiensis    波密无心菜
96(94)Petals shorter than sepals.(97)
+Petals equaling or longer than sepals.(98)
97(96)Cauline leaf blade ovate, petiole 0.5--1.5 cm; sepals white villous abaxially, margin membranous; petals ca. 2/3 as long as sepals, apex entire; styles 254  A. longipes    长梗无心菜
+Cauline leaf blade lanceolate, petiole absent or inconspicuous; sepals hairy abaxially and at margin; petals shorter than sepals, apex entire, sometimes 2-cleft; styles 2 or 355  A. inconspicua    不显无心菜
98(96)Petals 1.5--2 × as long as sepals; sepals leathery56  A. amdoensis    安多无心菜
+Petals equaling or up to 1.5 × as long as sepals; sepals herbaceous.(99)
99(98)Roots conic, fleshy; leaf blade linear or saccate.(100)
+Roots slender, fibrous, or with a rhizome; leaf blade linear-lanceolate or elliptic.(101)
100(99)Leaf blade linear; sepals glabrous, with wide membranous margin; petal apex entire58  A. membranisepala    膜萼无心菜
+Leaf blade saccate; sepals densely glandular pubescent abaxially; petal apex slightly emarginate or shallowly lobed57  A. spathulifolia    匙叶无心菜
101(99)Leaf blade linear-lanceolate, herbaceous, margin ciliate; petal apex erose59  A. karakorensis    克拉克无心菜
+Leaf blade elliptic, leathery, margin glabrous; petal apex entire60  A. tumengelaensis    土门无心菜
   List of lower taxa