Flora of China
  FOC Vol.8 (2001)              | Family List  Crassulaceae  PDF     | Link Font:+ Big | - Small
   Crassulaceae  Candolle
景天科   jing tian ke
Authors:Fu Kunjun (Fu Kun-tsun); Hideaki Ohba


Herbs, subshrubs, or shrubs. Stems mostly fleshy. Leaves alternate, opposite, or verticillate, usually simple; stipules absent; leaf blade entire or slightly incised, rarely lobed or imparipinnate. Inflorescences terminal or axillary, cymose, corymbiform, spiculate, racemose, paniculate, or sometimes reduced to a solitary flower. Flowers usually bisexual, sometimes unisexual in Rhodiola (when plants dioecious or rarely gynodioecious), actinomorphic, (3 or)4-6(-30)-merous. Sepals almost free or basally connate, persistent. Petals free or connate. Stamens as many as petals in 1 series or 2 × as many in 2 series. Nectar scales at or near base of carpels. Follicles sometimes fewer than sepals, free or basally connate, erect or spreading, membranous or leathery, 1- to many seeded. Seeds small; endosperm scanty or not developed.

About 35 genera and over 1500 species: Africa, America, Asia, Europe; 13 genera (two endemic, one introduced) and 233 species (129 endemic, one introduced) in China.

Some species of Crassulaceae are cultivated as ornamentals and/or used medicinally.

Fu Shu-hsia & Fu Kun-tsun. 1984. Crassulaceae. In: Fu Shu-hsia & Fu Kun-tsun, eds., Fl. Reipubl. Popularis Sin. 34(1): 31-220.

1Stamens in 1 series, usually as many as petals; flowers always bisexual.(2)
+Stamens usually in 2 series, 2 × as many as petals; flowers sometimes unisexual and male flowers without stamens.(5)
2(1)Leaves always opposite, joined to form a basal sheath; inflorescences axillary, often shorter than subtending leaf; plants not developing enlarged rootstock1  Tillaea    东爪草属
+Leaves alternate, occasionally opposite proximally; inflorescence terminal, often very large; plants sometimes developing enlarged, perennial rootstock.(3)
3(2)Inflorescence with a distinct, elongated main axis forming a raceme or thyrse much longer than broad; carpels narrowed at base, free; basal leaves often densely hairy4  Kungia    孔岩草属
+Inflorescence cymose with no distinct main axis, often broader than long; carpels broad at base, often ± connate; basal leaves often glabrous.(4)
4(3)Plants with conspicuous basal leaf rosettes; corolla campanulate, petals initially erect then spreading above middle, ± S-shaped in longitudinal section8  Sinocrassula    石莲属
+Plants without basal leaf rosettes, often ephemeral; petals rotate12  Sedum    景天属
5(1)Flowers 4-merous and with prominent, persistent corolla tube completely enclosing follicles; leaves opposite.(6)
+Flowers 5- or 6(-12)-merous, rarely 3- or 4-merous (Rhodiola) and then petals free or joined at base only and not enclosing follicles; leaves alternate, opposite, or in a rosette.(7)
6(5)Filaments inserted basally on corolla tube; flowers usually pendulous2  Bryophyllum    落地生根属
+Filaments inserted near or above middle of corolla tube; flowers erect3  Kalanchoe    伽蓝菜属
7(5)Carpels stipitate to substipitate or basally constricted or conspicuously attenuate, free.(8)
+Carpels sessile, base not constricted (slightly attenuate in some Rhodiola species) and usually connate (free in Sedum chuhsingense, S. semilunatum, and sometimes in S. przewalskii).(9)
8(7)Plants perennial, usually with several to many flowering stems; basal leaves not in a conspicuous rosette; inflorescence formed from a terminal group of cymes, usually broader than long, sometimes with aditional cymes in axils of distal leaves6  Hylotelephium    八宝属
+Plants monocarpic, with a single flowering stem; basal leaves in a somewhat conspicuous rosette, dying when plant flowers; inflorescence with an elongated main axis and numerous lateral cymules, sometimes a true raceme with cymules reduced to single flowe5  Orostachys    瓦松属
9(7)Stems dimorphic with usually very stout caudex or rhizome, usually with brown or blackish, membranous, scalelike leaves, sharply differentiated from much more slender, erect or ascending, leafy flowering stems.(10)
+Stems not sharply dimorphic, proximally with well developed leaves, rarely plants stoloniferous.(11)
10(9)Flowers bisexual; petals basally connate7  Pseudosedum    合景天属
+Flowers unisexual or bisexual; petals free or nearly so13  Rhodiola    红景天属
11(9)Leaf rosette absent at anthesis; inflorescence terminal.(12)
+Leaf rosette conspicuous at anthesis; inflorescence axillary.(13)
12(11)Leaves flattened, margin serrate or crenate; seed testa longitudinally costate or subsmooth11  Phedimus    费菜属
+Leaves terete or semiterete in cross section, margin entire (4-8-dentate in Sedum rosthornianum and remotely dentate in S. engleri var. dentatum); seed testa reticulate or papillate-reticulate12  Sedum    景天属
13(11)Petals yellow; basal leaf rosette almost as wide as inflorescences are long; plants glabrous throughout10  Ohbaea    岷江景天属
+Petals white, pink, and/or red, (rarely pale yellow outside China); plants hairy or glabrous.(14)
14(13)Plants glandular hairy or, if glabrous, flowers 6-8-merous; petals connate at base9  Rosularia    瓦莲属
+Plants glabrous; flowers (3-)5-merous; petals free or almost so13  Rhodiola    红景天属
   List of lower taxa