Flora of China
  FOC Vol.19 (2011)              | Family List  Cucurbitaceae  PDF Thladiantha  PDF   | Link Font:+ Big | - Small
   11. Thladiantha  Bunge Enum. Pl. Chin. Bor. 29. 1833.
赤瓟属   chi bao shu
Authors:Authors: Anmin Lu & Charles Jeffrey


Herbs, perennial, climbing, scandent, or prostrate. Roots usually tuberous. Stems herbaceous, scabrous. Tendrils simple or 2-fid. Leaves mostly simple, vary rarely pedately 3-7-foliolate; leaf blade cordate, margin dentate. Plants dioecious. Male flowers in cymes-racemes or rarely solitary; calyx tube broadly campanulate or cupular; segments 5, linear, lanceolate, ovate-lanceolate, or oblong, 1-3-veined; corolla yellow, campanulate, 5-partite; segments oblong, broadly ovate, or obovate, usually 5-7-veined, entire; stamens 5, inserted on calyx tube, free, often 4 in pairs, other 1 free; filaments short; anthers oblong or ovoid, all 1-celled; anther cell erect; rudimentary ovary glandlike. Female flowers solitary or 2- or 3-fascicled on a short peduncle; calyx and corolla as in male flowers; ovary ovoid, oblong, or fusiform, smooth or verrucose; ovules numerous, horizontal; style 3-fid; stigmas 3, 2-lobed, reniform. Fruit fleshy, indehiscent, smooth or verrucose, many ribbed or not ribbed. Seeds numerous, horizontal.

Twenty-three species: Bhutan, China, India, Indonesia, Japan (introduced), Korea, Laos, Myanmar, Nepal, Thailand, Vietnam; 23 species (19 endemic) in China.

1Bracts in male flowers flabelliform, dentate(2)
+Male flowers without flabelliform bracts(4)
2(1)Calyx segments entire, 0.6-3 mm wide.1  T. cordifolia    大苞赤瓟
+Calyx segments dentate or irregularly lobed, (3-)4-11 mm wide(3)
3(2)Calyx segments (3-)4-6 mm wide, with 1-3 triangular lobes.2  T. grandisepala    大萼赤瓟
+Calyx segments to 11 mm wide, flabelliform, margin irregularly lobed.3  T. medogensis    墨脱赤瓟
4(1)Ovary and fruit scaly or verrucose(5)
+Ovary and fruit not scaly or verrucose(7)
5(4)Leaf blade ovate-lanceolate or narrowly ovate-triangular; ovary and fruit verrucose.23  T. longifolia    长叶赤瓟
+Leaf blade ovate-cordate; fruit with thick scales(6)
6(5)Ovary and fruit ovoid-globose, apex with a long rostrum, scales at base of fruit less than 5 mm, not adnate to pedicel.21  T. pustulata    云南赤瓟
+Ovary and fruit ovoid-oblong, apex without a rostrum, scales at base of fruit to 10 mm, adnate to pedicel.22  T. henryi    皱果赤瓟
7(4)Leaves palmately deeply 5-lobed.20  T. palmatipartita    掌叶赤瓟
+Leaves not palmately deeply 5-lobed(8)
8(7)Male flowers congested at apex of rachis into pseudocapitulum(9)
+Male flowers in cyme-panicle, panicle, raceme, or rarely solitary, but not capitellate(10)
9(8)Calyx segments lanceolate, apex acute, 1-veined.18  T. capitata    头花赤瓟
+Calyx segments linear, apex obtuse, 3-veined.19  T. davidii    川赤瓟
10(8)Plants sparsely hairy or subglabrous(11)
+Plants with dense multicellular setae, very rarely subglabrous(15)
11(10)Calyx segments 1-veined(12)
+Calyx segments 3-veined(13)
12(11)Basal lateral vein along base of leaf blade; calyx segments linear, over 9 mm.13  T. oliveri    鄂赤瓟
+Basal lateral vein not along base of leaf blade; calyx segments triangular- lanceolate, 4-5 mm.14  T. montana    山地赤瓟
13(11)Tendrils 2-fid, leaves not lobed or lobed.17  T. dentata    齿叶赤瓟
+Tendrils simple, leaves not lobed(14)
14(13)Leaf blade broadly ovate-cordate; tendrils filiform; pedicel in male flowers 1.2-2.5 cm; fruit with a long beak.15  T. maculata    斑赤瓟
+Leaf blade narrowly ovate or narrowly ovate-lanceolate; tendrils robust; pedicel 0.5-1 cm; fruit without beak.16  T. punctata    台湾赤瓟
15(10)Leaves simple, unlobed or margin lobed or serrate, or pedately 3-5-foliolate.4  T. hookeri    异叶赤瓟
+Leaves unlobed, rarely irregularly lobed(16)
16(15)Tendrils simple(17)
+Tendrils 2-fid(19)
17(16)Male flowers solitary or in a short raceme, bracts leaflike.5  T. dubia    赤瓟
+Male flowers without bracts(18)
18(17)Petiole less than 1.5 cm, leaf blade lanceolate or narrowly ovate.6  T. sessilifolia    短柄赤瓟
+Petiole over 2 cm, leaf blade ovate-cordate.7  T. villosula    长毛赤瓟
19(16)Male flowers dimorphic, solitary, and in raceme with (2 or)3(-5) flowers in axil of same leaf.8  T. dimorphantha    山西赤瓟
+Male flowers in panicle or raceme(20)
20(19)Stem and leaves with pale yellow multicellular setae or very rarely subglabrous, sinus at base of leaf closed; fruit orange-red when mature.9  T. nudiflora    南赤瓟
+Stem and leaves with gray-white multicellular setae, sinus at base of leaf open; fruit whitish yellow when mature(21)
21(20)Calyx segments broadly lanceolate or obovate-oblong, 10-15 mm.10  T. longisepala    长萼赤瓟
+Calyx segments triangular-lanceolate, less than 10 mm(22)
22(21)Ovary and fruit with dense yellow-brown spiniform setae; seeds 6.5-7 mm, sparsely irregularly verrucose on surface.11  T. setispina    刚毛赤瓟
+Ovary and fruit with long setae; seeds 4.5-5 mm, not verrucose on surface.12  T. lijiangensis    丽江赤瓟
   List of lower taxa