foc
《中国植物志》英文修订版
Flora of China
  FOC Vol.19 (2011)              | Family List  Morinaceae  PDF Morina  PDF   |  eFloras.org Link Font:+ Big | - Small
   2. Morina  Linnaeus Sp. Pl. 1: 28. 1753.
刺参属   ci shen shu
  

Asaphes Sprengel (1827), not Candolle (1825); Cryptothladia (Bunge) M. J. Cannon; Morina sect. Cryptothladia Bunge.

  

Roots usually thickened, branched. Caudex short, woody, usually covered with remnants of leaf bases. Leaves in whorls of 3 or 4(-6), rarely 2, opposite, linear to oblong-lanceolate, entire to pinnatipartite, spinose. Inflorescence of several verticillasters, each subtended by a whorl of leaflike bracts. Flowers sessile or shortly pedicellate, enveloped in an involucel; involucels campanulate, with 8-16 spiny teeth, 2 significantly longer than others. Calyx tube oblique, campanulate; limb 2-lipped, lips 2- or 3-lobed or emarginate. Corolla tube elongate; limb spreading, 2-lipped; upper lip 2-lobed, lower one 3-lobed. Fertile stamens 2, inserted at corolla throat; staminodes 2, at base of corolla tube, cordate. Nectary 1, anterior, at base of corolla tube, 3-lobed. Ovary inferior, 1-loculed, enveloped in an involucel; style usually longer than stamens; stigma disklike; ovule single, pendulous. Achenes rugose, columnar, slightly or markedly obliquely truncate.

About ten species: from the Balkans to C Asia and E Himalaya; eight species (four endemic) in China.

Cannon and Cannon (Bull. Brit. Mus. (Nat. Hist.), Bot. 12(1): 1-35. 1984) treated Cryptothladia at the rank of genus, separating it from Morina. Hong (Novon 20: 418-419. 2010) has explained why this taxon would be better merged into the genus Morina.

Cannon and Cannon (Bull. Brit. Mus. (Nat. Hist.), Bot. 12(1): 1-35. 1984) treated Cryptothladia at the rank of genus, separating it from Morina. Hong (Novon 20: 418-419. 2010) has explained why this taxon would be better merged into the genus Morina.

  
1Involucral bracts connate, forming shallow cups; involucel teeth much shorter than tube (1/2 or less as long as tube).6  M. polyphylla    多叶刺参
+Involucral bracts free; involucel teeth at least 2/3 as long as tube(2)
   
2(1)Leaves with petioles 4-8 cm; leaf margins spinose, rarely sinuate-dentate(3)
+Leaves sessile or subsessile; leaf margins mostly lobed, less frequently entire(4)
   
3(2)Involucral bracts acuminate at apex, spinose along whole margins; calyx lips 2-lobed, lobes lanceolate, 5-6 mm; middle and upper cauline leaves petiolate.1  M. chlorantha    绿花刺参
+Involucral bracts acute or rounded at apex, spinose only at base; calyx emarginate; middle and upper cauline leaves subsessile.2  M. bracteata    宽苞刺参
   
4(2)Leaves entire or slightly toothed; flowers in 20 or more closely packed whorls of 8-12.5  M. ludlowii    藏南刺参
+Leaves lobed; flowers in fewer, usually interrupted, whorls(5)
   
5(4)Corolla 2-3 cm, greatly exceeding calyx, obviously 5-lobed; staminodes inserted at middle part of corolla tube(6)
+Corolla less than 1 cm, almost hidden by calyx, weakly lobed with 2 or 4 major lobes; staminodes inserted at base of corolla tube(7)
   
6(5)Involucral bracts broadly ovate or orbicular, ca. as long as wide (excluding pointed apex); calyx covered by involucral bracts.7  M. longifolia    长叶刺参
+Involucral bracts broadly ovate-lanceolate, much longer than wide; calyx exposed.8  M. coulteriana    黄花刺参
   
7(5)Calyx lobes 4, ovate, apex rounded; leaves shallowly lobed.3  M. chinensis    刺参
+Calyx lobes 4-6, narrowly ovate or ovate-lanceolate, apex usually spinose; leaves cleft almost to midvein.4  M. kokonorica    青海刺参
   
   Lower Taxon
  • Morina chinensis  Y. Y. Pai  刺参
  • Morina chlorantha  Diels  绿花刺参
  • Morina kokonorica  K. S. Hao  青海刺参
  • Morina bracteata  C. Y. Cheng & H. B. Chen  宽苞刺参
  • Morina longifolia  Wallich ex Candolle  长叶刺参
  • Morina polyphylla  Wallich ex Candolle  多叶刺参
  • Morina coulteriana  Royle  黄花刺参
  • Morina ludlowii  (M. J. Cannon) D. Y. Hong  藏南刺参