Flora of China
  FOC Vol.8 (2001)              | Family List  Brassicaceae  PDF Lepidium  PDF   | Link Font:+ Big | - Small
   10. Lepidium  Linnaeus
独行菜属   du xing cai shu


Herbs annual, biennial, or perennial, sometimes subshrubs, rarely shrubs or climbers. Trichomes absent or simple. Stems erect or ascending, sometimes creeping, simple or branched basally and/or apically. Basal leaves rosulate or not, simple, entire or pinnately dissected. Cauline leaves petiolate or sessile, base cuneate, attenuate, auriculate, sagittate, or amplexicaul, margin entire, dentate, or dissected. Racemes ebracteate, corymbose, elongated or not in fruit. Fruiting pedicels terete, flattened, or winged, erect or divaricate. Sepals ovate or oblong, rarely orbicular, base of lateral pair not saccate. Petals white, yellow, or pink, erect or spreading, sometimes rudimentary or absent; blade obovate, spatulate, oblong, oblanceolate, orbicular, linear, or filiform, apex obtuse, rounded, or emarginate; claw absent or distinct. Stamens 2 and median, sometimes 6 and tetradynamous or subequal in length, rarely 4 and all median or 2 median and 2 lateral; anthers ovate or oblong. Nectar glands 4 or 6, distinct; median glands always present. Ovules 2 per ovary; placentation apical. Fruit dehiscent silicles, oblong, ovate, obovate, cordate, obcordate, elliptic, or orbicular, strongly angustiseptate; valves veinless or prominently veined, keeled or not, apically winged or wingless; replum rounded; septum complete or perforated, membranous; style absent, obsolete, or distinct, included or exserted from apical notch of fruit; stigma capitate, entire or rarely 2-lobed. Seeds 1 per locule, winged, margined, or wingless, oblong or ovate, plump or flattened; seed coat smooth, minutely reticulate, or papillate, usually copiously mucilaginous when wetted; cotyledons incumbent, rarely accumbent or diplecolobal.

About 180 species: all continents except Antarctica; 16 species (two endemic, two introduced) in China.

1Cauline leaves sessile, amplexicaul, sagittate, or auriculate at base.(2)
+Cauline leaves petiolate or subsessile, cuneate or obtuse at base, if subauriculate then plants with capitate or clavate papillae.(5)
2(1)Flowers yellow; upper leaves amplexicaul-perfoliate, basal leaves 2- or 3-pinnatisect; seeds winged all around10  L. perfoliatum    抱茎独行菜
+Flowers white; upper leaves auriculate or sagittate, basal leaves entire, dentate, or pinnatifid; seeds wingless.(3)
3(2)Perennials, glabrous, papillate, or crisped hairy; fruit wings free from style; leaves leathery.(4)
+Annuals or biennials, usually densely long pubescent; fruit wings united with lower part of style; leaves herbaceous1  L. campestre    绿独行菜
4(3)Basal and lowermost cauline leaves pinnatifid, upper ones subentire; plants glabrous3  L. cordatum    心叶独行菜
+All leaves entire; plants with curved papillate trichomes4  L. cartilagineum    碱独行菜
5(1)Stamens 6; stems glabrous or pubescent with long trichomes (puberulent in L. alashanicum).(6)
+Stamens 4 or 2; stems puberulent with clavate or capitate papillae, sometimes with minute trichomes.(11)
6(5)Petals absent; plants puberulent with subclavate papillate trichomes; all leaves linear, entire, margin involute9  L. alashanicum    阿拉善独行菜
+Petals well developed; plants glabrous or pilose with slender trichomes; leaves variously-shaped, at least some pinnatifid, pinnatisect, dentate, or serrate, margin flat.(7)
7(6)Annuals; fruit (4-)5-6 mm, winged all around, broadly so apically; all or at least lower leaves 1- or 2-pinnatisect or -pinnatifid; cotyledons 3-lobed2  L. sativum    家独行菜
+Perennials; fruit 1.5-3 mm, wingless or only apically winged; leaves entire, serrate, or dentate; cotyledons entire.(8)
8(7)Sepals persistent in fruit; fruit cordate at base7  L. obtusum    钝叶独行菜
+Sepals deciduous shortly after anthesis; fruit cuneate, obtuse, or rounded at base.(9)
9(8)Basal and lowermost cauline leaves pinnatifid or lyrate, with oblong or linear lateral lobes much smaller than terminal one; style (0.2-)0.3-0.4(-0.5) mm, distinctly exserted from apical notch8  L. lacerum    裂叶独行菜
+Basal and lowermost leaves dentate, serrate, or subentire; style absent, obsolete, or rarely to 0.1 mm.(10)
10(9)Uppermost cauline leaves sessile or subsessile, leathery; fruit not carinate; seeds (0.8-)1-1.3 mm6  L. latifolium    宽叶独行菜
+Uppermost cauline leaves shortly petiolate, papery; fruit carinate; seeds 1.5-2.2 mm5  L. ferganense    全缘独行菜
11(5)Stamens 4; style longer than and exserted from apical notch of fruit11  L. cuneiforme    楔叶独行菜
+Stamens 2; style shorter than and included within apical notch of fruit.(12)
12(11)Fruit orbicular; upper stems and rachis puberulent with curved minute trichomes; cotyledons accumbent13  L. virginicum    北美独行菜
+Fruit elliptic, ovate, or obovate; upper stems and rachis puberulent with straight or clavate papillae; cotyledon incumbent.(13)
13(12)Middle and upper cauline leaves usually pinnatifid; infructescences capitate; petals 0.5-1 mm12  L. capitatum    头花独行菜
+Middle and upper cauline leaves dentate, serrate, or entire; infructescences elongated racemes; petals absent or rudimentary.(14)
14(13)Basal leaves 2- or 3-pinnatisect; pedicels puberulent all around with slender trichomes; herbs fetid14  L. ruderale    柱毛独行菜
+Basal leaves 1-pinnatifid or serrate; pedicel papillate only adaxially; herbs not fetid.(15)
15(14)Fruit broadly obovate, widest above middle; raceme papillae terete or subclavate15  L. densiflorum    密花独行菜
+Fruit elliptic, widest at middle; raceme papillae clavate or capitate16  L. apetalum    独行菜
   List of lower taxa