Flora of China
  FOC Vol.8 (2001)              | Family List  Brassicaceae  PDF Isatis  PDF   | Link Font:+ Big | - Small
   13. Isatis  Linnaeus
菘蓝属   song lan shu


Herbs annual, biennial, or perennial, often glaucous. Trichomes absent or simple. Stems erect, simple at base, paniculately branched above. Basal leaves petiolate or rarely sessile, rosulate or not, simple, entire, dentate, or pinnately lobed. Cauline leaves sessile and auriculate, sagittate, or amplexicaul at base, rarely petiolate and attenuate, entire or dentate. Racemes many flowered, ebracteate, forming panicles, elongated considerably in fruit. Fruiting pedicels filiform, often thickened and clavate at apex, reflexed. Sepals ovate or oblong, erect or ascending, base of lateral pair not saccate. Petals yellow, as long as or longer than sepals; blade obovate, spatulate, oblong, or oblanceolate, apex obtuse or subemarginate; claw absent. Stamens 6, slightly tetradynamous; anthers ovate or oblong, apiculate or obtuse at apex. Nectar glands confluent, or 4 and median and lateral pairs distinct. Ovules 1(or 2) per ovary, subapical. Fruit indehiscent, samaroid siliques or silicles, oblong, ovate, obovate, cordate, elliptic, oblanceolate, spatulate, or orbicular, strongly angustiseptate, sessile, prominently winged all around or distally, 1(or 2)-seeded, glabrous or hairy, smooth; seed-bearing locule papery or corky, prominently or obscurely 1- or 3-veined, sometimes keeled or shortly winged; valves and replum united; gynophore, style, and septum absent; stigma capitate, entire. Seeds wingless, narrowly oblong, plump; seed coat smooth, not mucilaginous when wetted; cotyledons incumbent or accumbent.

About 50 species: primarily in C and SW Asia; four species in China.

Mature fruit are essential for the reliable determination of species. Leaf, flower, and indumentum characters are unreliable for the determination of Chinese and C Asian species.

1Fruit winged only above the middle, constricted and often curved at middle, spatulate, often with crisped trichomes; petals 1-2 mm4  I. minima    小果菘蓝
+Fruit winged all around, not curved and rarely slightly constricted at middle, variously-shaped but not spatulate, trichomes straight; petals (2.2-)2.5-4 mm.(2)
2(1)Fruiting pedicels slender throughout; fruit oblong-pandurate, densely puberulent with subclavate trichomes, apex emarginate or rarely truncate1  I. violascens    宽翅菘蓝
+Fruiting pedicels clavate at apex; fruit oblong, elliptic, oblong-oblanceolate, or elliptic-obovate, glabrous or pubescent with slender trichomes, apex acute, obtuse, rounded, or rarely subemarginate.(3)
3(2)Fruit only with a prominent midvein and inconspicuous lateral veins, widest above middle, black or blackish brown2  I. tinctoria    菘蓝
+Fruit with prominent midvein and two lateral veins, widest at middle, light brown3  I. costata    三肋菘蓝
   Lower Taxon
  • Isatis costata  C. A. Meyer  三肋菘蓝
  • Isatis minima  Bunge  小果菘蓝
  • Isatis tinctoria  Linnaeus  菘蓝
  • Isatis violascens  Bunge  宽翅菘蓝