Flora of China
  FOC Vol.6 (2001)              | Family List  Caryophyllaceae  PDF Gypsophila  PDF   | Link Font:+ Big | - Small
   29. Gypsophila  Linnaeus
石头花属   shi tou hua shu
Authors:Lu Dequan; Nicholas J. Turland

Heterochroa Bunge; Psammophila Fourreau; Psammophiliella Ikonnikov; Timaeosia Klotzsch.


Herbs annual or perennial. Stems usually caespitose, erect or diffuse, glabrous or glandular pubescent, sometimes glabrescent or base woody. Leaves opposite, linear, lanceolate, ovate, oblong, spatulate, or sometimes subulate or subfleshy. Flowers numerous, bisexual, small, borne in dichasial cymes, corymbs, or panicles, sometimes aggregated into capitula; bracts usually scarious, rarely leaflike. Calyx green or purple veined, white membranous between veins, campanulate or funnelform, rarely cylindric, glabrous or puberulous, veins broad, apex 5-toothed. Petals 5, white or pink, sometimes purple veined, oblong or obovate, longer than calyx, base usually cuneate, apex entire, truncate, or retuse. Stamens 10. Ovary globose or ovoid, 1-loculed; ovules numerous. Styles 2(or 3). Gynophore absent. Capsule globose, ovoid, or oblong, 4-valved. Seeds subreniform, compressed, tuberculate; hilum lateral; embryo annular; radicle prominent.

About 150 species: mainly in temperate Asia and Europe, fewer species in NE Africa (Egypt), Australia, and North America; 17 species (four endemic) in China.

Gypsophila elegans Marschall von Bieberstein, native to SW Asia and SE Europe, is cultivated in China as an ornamental.

1Herbs annual17  G. muralis    细小石头花
+Herbs perennial.(2)
2(1)Stems densely caespitose, ascending; bracts leaflike; calyx not white membranous between veins.(3)
+Stems simple or laxly caespitose, erect; bracts scarious; calyx white membranous between veins.(5)
3(2)Plants white pubescent; leaves obovate-spatulate14  G. cerastioides    卷耳状石头花
+Plants brown glandular pubescent or sericeous; leaves linear-subulate or lanceolate.(4)
4(3)Plants brown glandular pubescent; leaves linear-subulate; pedicels 0.3--1.2 cm15  G. desertorum    荒漠石头花
+Plants sericeous; leaves lanceolate; pedicels 1.4--2 cm16  G. sericea    绢毛石头花
5(2)Inflorescence paniculate, spreading, many branched, pedicels slender; calyx broadly campanulate.(6)
+Inflorescence usually corymbose, sometimes subcapitate, pedicels usually stout, rigid; calyx campanulate or narrowly so.(7)
6(5)Leaves lanceolate or linear-lanceolate, 0.25--0.7 cm wide, glabrous; calyx 1.5--2 mm, veins purple12  G. paniculata    圆锥石头花
+Leaves obovate- or ovate-oblong, 1--3 cm wide, glandular pubescent; calyx 2--4 mm, veins green13  G. perfoliata    钝叶石头花
7(5)Leaves linear.(8)
+Leaves not linear.(11)
8(7)Inflorescence few branched or subcapitate, few flowered.(9)
+Inflorescence many branched, numerous flowered.(10)
9(8)Inflorescence lax, pedicels 5--20 mm; calyx 2--3 mm8  G. patrinii    紫萼石头花
+Inflorescence subcapitate, pedicels ca. 1 mm; calyx 3.5--5 mm9  G. capituliflora    头状石头花
10(8)Inflorescence lax, divergent spinose11  G. spinosa    刺序石头花
+Inflorescence congested, not spinose10  G. licentiana    细叶石头花
11(7)Leaves linear-lanceolate.(12)
+Leaves linear-oblanceolate, oblong, or ovate.(14)
12(11)Plants more than 50 cm tall; inflorescence many branched; calyx veins green6  G. davurica    草原石头花
+Plants less than 50 cm tall; inflorescence few branched; calyx veins purple-brown.(13)
13(12)Inflorescence lax, few flowered, pedicels 5--15 mm; calyx 3--5 mm, lobes acuminate; stamens and styles included; seeds obtusely tuberculate4  G. tschiliensis    河北石头花
+Inflorescence subcapitate, numerous flowered, pedicels 2--3 mm; calyx 2--3 mm, lobes obtuse; stamens and styles exserted; seeds acutely tuberculate5  G. huashanensis    华山石头花
14(11)Leaves linear-oblanceolate.(15)
+Leaves oblong or ovate.(16)
15(14)Plants 50--80 cm tall; inflorescences scattered, pedicels 2--5 mm; bracts and calyx lobes ciliate at margin; seeds acutely tuberculate1  G. altissima    高石头花
+Plants 20--50 cm tall; inflorescences concentrated into capitula, pedicels 0.5--2.5 mm; bracts and calyx lobes undulate at margin, not ciliate; seeds truncately tuberculate2  G. cephalotes    膜苞石头花
16(14)Leaves oblong; inflorescence dense, pedicels 2--5 mm; petal apex truncate or retuse; stamens and styles longer than petals3  G. oldhamiana    长蕊石头花
+Leaves ovate; inflorescence spreading, pedicels 5--10 mm; petal apex rounded; stamens and styles shorter than petals7  G. pacifica    大叶石头花
   List of lower taxa