Flora of China
  FOC Vol.8 (2001)              | Family List  Brassicaceae  PDF Eutrema  PDF   | Link Font:+ Big | - Small
   89. Eutrema  R. Brown
山(艹俞)菜属   shan yu cai shu

Wasabia Matsumura.


Herbs perennial, rhizomatous or with a caudex. Trichomes absent or simple. Stems erect, ascending, or decumbent, simple or branched apically. Basal leaves petiolate, rosulate, simple, entire or palmately lobed, palmately veined. Cauline leaves petiolate or sessile and cuneate or auriculate, pinnately or palmately veined, entire, dentate, or crenate; ultimate veins ending or not with apiculate callosities. Racemes ebracteate or rarely bracteate throughout, elongated considerably or not elongated in fruit. Sepals ovate or oblong, base of lateral pair not saccate. Petals white or rarely pink; blade spatulate or rarely obovate, apex obtuse; claw absent. Stamens 6, slightly tetradynamous; filaments slightly dilated at base; anthers ovate or oblong, obtuse at apex. Nectar glands confluent and subtending bases of all stamens; median glands present. Ovules 2-10 per ovary. Fruit dehiscent, siliques or silicles, linear, oblong, ovoid, or lanceolate, terete or slightly 4-angled, sessile or shortly stipitate; valves with an obscure or prominent midvein, smooth or torulose; replum rounded; septum complete or perforated, translucent, veinless; style obsolete or distinct and to 3 mm; stigma capitate, entire. Seeds uniseriate, wingless, oblong, plump; seed coat obscurely reticulate, not mucilaginous when wetted; cotyledons incumbent.

Nine species: primarily C and E Asia and Himalayas, one species extending into North America; seven species (one endemic, one introduced) in China.

1Middle and upper cauline leaves sessile, pinnately veined.(2)
+All leaves petiolate, palmately veined.(4)
2(1)Main infructescence 0.2-2(-3.5) cm, often subumbellate; plants 2-15(-25) cm tall; petals 2-3.5 mm; sepals often persistent to fruit maturity1  E. heterophyllum    密序山(艹俞)菜
+Main infructescence 10-20(-40) cm tall, lax racemes; plants 30-110 cm tall; petals 4-5 mm; sepals caducous.(3)
3(2)Plants glabrous throughout; uppermost leaves cuneate at base; fruit 5-8(-10) mm2  E. integrifolium    全缘叶山(艹俞)菜
+Plants sparsely to densely pilose at least along midvein and leaf margin; uppermost leaves auriculate or amplexicaul at base; fruit (8-)12-20(-25) mm3  E. himalaicum    川滇山(艹俞)菜
4(1)Fruiting pedicels erect or ascending; ultimate leaf veins not ending in apiculate callosities; fruit lanceolate, ovoid, or oblong, not torulose, often subappressed to rachis, slightly 4-angled4  E. deltoideum    三角叶山(艹俞)菜
+Fruiting pedicels divaricate or reflexed; ultimate leaf veins ending in apiculate callosities; fruit usually linear, torulose, not appressed to rachis, terete.(5)
5(4)Rhizomes fleshy; petals 6-8(-9) mm; sepals 3-4 mm; gynophore (1-)2-5 mm; filaments 3.5-5 mm7  E. wasabi    块茎山(艹俞)菜
+Rhizomes not fleshy; petals 3.5-5(-6) mm; sepals 1.5-2 mm; gynophore absent or obsolete; filaments 1.5-2 mm.(6)
6(5)Racemes bracteate throughout6  E. tenue    日本山(艹俞)菜
+Racemes ebracteate or only lowermost few flowers bracteate5  E. yunnanense    南山(艹俞)菜
   Lower Taxon
  • Eutrema deltoideum  (J. D. Hooker & Thomson) O. E. Schulz  三角叶山(艹俞)菜
  • Eutrema integrifolium  (de Candolle) Bunge  全缘叶山(艹俞)菜
  • Eutrema tenue  (Miquel) Makino  日本山(艹俞)菜
  • Eutrema yunnanense  Franchet  南山(艹俞)菜
  • Eutrema heterophyllum  (W. W. Smith) H. Hara  密序山(艹俞)菜
  • Eutrema himalaicum  J. D. Hooker & Thomson  川滇山(艹俞)菜
  • Eutrema wasabi  (Siebold) Maximowicz  块茎山(艹俞)菜
  • Eutrema bouffordii  Al-Shehbaz