Flora of China
  FOC Vol.6 (2001)              | Family List  Caryophyllaceae  PDF Cerastium  PDF   | Link Font:+ Big | - Small
   13. Cerastium  Linnaeus
卷耳属   juan er shu
Authors:Lu Dequan; John K. Morton


Herbs annual or perennial, pubescent and/or glandular pubescent, rarely glabrous. Stems usually caespitose, sometimes slightly woody at base. Leaves ovate, elliptic, or lanceolate. Inflorescence terminal, a dichasial cyme, dense or lax, sometimes flowers solitary. Sepals (4 or)5, free. Petals (4 or)5, sometimes absent, white, apex usually 2-lobed or retuse, rarely entire. Stamens (3 or 5 or)10; nectaries present. Ovary 1-loculed; ovules numerous. Styles 3--5, inserted opposite sepals. Capsule golden yellowish, cylindric, sometimes subequaling but usually exceeding calyx, hard, thin, brittle, dehiscing by 2 × as many teeth as styles; teeth slightly curved, sometimes straight or revolute. Seeds numerous, globose or reniform, compressed, usually tuberculate; embryo annular.

About 100 species: almost cosmopolitan, most abundant in temperate and cold regions: 23 species (nine endemic) in China.

The identities of the taxa to which the following three names apply could not be ascertained. All were described from China, but none of the types has been seen by the present authors: Cerastium calcicola Ohwi (Repert. Spec. Nov. Regni Veg. 36: 46. 1934), described from Taiwan; C. kaoi Shimizu (J. Fac. Text. Sci. & Techn. Shinshu Univ. No. 36, Biol., No. 12 [Stud. Limest. Fl. Jap. & Taiwan, Pt. 2]: 23. 1963), also described from Taiwan; and C. sinicum Nakai (J. Jap. Bot. 15: 526. 1939), described from Hebei.

The record of Cerastium perfoliatum Linnaeus in FRPS (p. 83), from Zhejiang, requires confirmation. No specimens of this species have been seen by the current authors.

1Styles 3; capsule teeth 6.(2)
+Styles (4 or)5; capsule teeth (8 or)10.(3)
2(1)Leaves linear-lanceolate; petals obovate, longer than sepals; stamens 101  C. cerastoides    六齿卷耳
+Leaves ovate-elliptic; petals ovate, shorter than sepals; stamens 52  C. parvipetalum    小瓣卷耳
3(1)Plants up to 20 cm tall.(4)
+Plants at least 15 cm tall.(12)
4(3)Leaves elliptic, oblong, ovate, or obovate-elliptic.(5)
+Leaves linear-lanceolate, lanceolate, or ovate-lanceolate.(8)
5(4)Cyme densely clustered, headlike3  C. glomeratum    球序卷耳
+Cyme not as above.(6)
6(5)Stems ascending; pedicel 5--8 mm, usually nutant after anthesis4  C. pusillum    山卷耳
+Stems erect; pedicel to 15 mm.(7)
7(6)Leaf apex obtuse; flowers sparse; sepals oblong, 4--5 mm5  C. limprichtii    椭圆叶卷耳
+Leaf apex acute; flowers dense; sepals oblong-lanceolate, 7--8 mm6  C. thomsonii    藏南卷耳
8(4)Leaves ovate-lanceolate.(9)
+Leaves linear-lanceolate or lanceolate.(10)
9(8)Plants annual; leaves 1--3 cm × 2--5 mm; flowers borne in a lax dichasium; calyx strongly inflated in fruit; capsule conical-cylindric7  C. dichotomum    膨萼卷耳
+Plants perennial; leaves 1--2.5 cm × 3--10 mm; flower solitary; calyx not inflated in fruit; capsule cylindric8  C. lithospermifolium    紫草叶卷耳
10(8)Leaf apex obtuse; petals slightly shorter than or equaling sepals9  C. szechuense    四川卷耳
+Leaves apex acute; petals longer than sepals.(11)
11(10)Leaves 1.3--2 cm × 2--4 mm; petals ca. 0.5 cm10  C. baischanense    长白卷耳
+Leaves 0.5--1 cm × 1--2 mm; petals 1--1.3 cm11  C. morrisonense    玉山卷耳
12(3)Leaves ovate, oblong, or elliptic.(13)
+Leaves linear-lanceolate, lanceolate, ovate-lanceolate, or oblanceolate.(16)
13(12)Plants stout, 40--100 cm tall; leaves 5--8 × 1.5--4 cm.(14)
+Plants slender, 15--60 cm tall; leaves 1--4 × 0.3--1.5 cm.(15)
14(13)Leaves and bracts opposite12  C. davuricum    达乌里卷耳
+Leaves and bracts 3-verticillate13  C. verticifolium    轮叶卷耳
15(13)Stems glabrescent; petals ca. 2 × as long as sepals14  C. wilsonii    卵叶卷耳
+Stems pilose and/or glandular pubescent; petals slightly shorter than or equaling sepals15  C. fontanum    喜泉卷耳
16(12)Leave ovate-lanceolate or oblanceolate.(17)
+Leaves lanceolate or linear-lanceolate.(18)
17(16)Plants perennial; leaves ovate-lanceolate; sepals oblong-lanceolate, ca. 5 mm; petals oblong or obovate, apex bifid16  C. furcatum    缘毛卷耳
+Plants annual; leaves oblanceolate; sepals ovate, 2--3 mm; petals spatulate, apex entire17  C. takasagomontanum    高山卷耳
18(16)Leaves lanceolate.(19)
+Leaves linear-lanceolate.(20)
19(18)Leaves 0.2--1 cm wide; sepals lanceolate; petal apex 2-lobed18  C. falcatum    披针叶卷耳
+Leaves 1--2 cm wide; sepals ovate-oblong; petal apex entire19  C. pauciflorum    疏花卷耳
20(18)Leaves 1--2.5 cm; petals apically 2-lobed for 1/4--1/3 their length20  C. arvense
+Leaves 3--8 cm; petals apically entire, retuse, or crenately 2-lobed.(21)
21(20)Petals spatulate, apex entire21  C. subpilosum    毛卷耳
+Petals obovate, apex retuse or crenately 2-lobed.(22)
22(21)Leaves 3--5 cm × 3--6 mm; sepals lanceolate; petals slightly longer than sepals; capsule ca. 2 × as long as sepals22  C. tianschanicum    天山卷耳
+Leaves 4--8 cm × 5--15 mm; sepals ovate; petals 2--3 × as long as sepals; capsule 2.5--3 × as long as sepals23  C. maximum    大卷耳
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